Pyrotherium

Name: Pyrotherium ‭(‬Fire beast‭)‬.
Phonetic: Py-roe-fee-re-um.
Named By: Florentino Ameghino‭ ‬-‭ ‬1888.
Classification: Chordata,‭ ‬Mammalia,‭ ‬Pyrotheria,‭ ‬Pyrotheriidae.
Species: P.‭ ‬romeroi‭ (‬type‭)‬,‭ ‬P.‭ ‬macfaddeni.‭ ‬Additionally,‭ ‬P.‭ ‬giganteum,‭ ‬P.‭ ‬planum,‭ ‬P.‭ ‬sorondoi.
Diet: Herbivore.
Size: Skull up to 72 centimetres long. Bodly length between 3 and 3.5 meters.
Known locations: South America.
Time period: Rupelian of the Oligocene.
Fossil representation: Several specimens.

       Pyrotherium was a fairly large and heavily built quadrupedal mammal that seems to have been similar to primitive elephants that were living in other parts of the world.‭ ‬This is yet another case of convergent evolution that is displayed by South American mammals which during this time would have been living upon a continent that was isolated from the rest of the world.‭ ‬Other examples‭ ‬include‭ ‬later litopterns such a Theosodon which resembled a primitive horse and Macrauchenia which looked like a bizarre camel.
       Pyrotherium possesses highly modified incisors‭ ‬that‭ ‬grew into short tusks that extended from both upper‭ (‬two pairs‭) ‬and lower jaws‭ (‬one pair‭)‬.‭ ‬The anterior portion of the skull is also shaped so that a short trunk grew from the end of the snout.‭ ‬Together Pyrotherium would have used the tusks to root up plants and then manipulate them into its mouth with the trunk.‭ ‬Once inside the molar teeth at the back of the mouth would process the food before it was swallowed.

Further reading
- Sur les oiseaux fossiles de Patagonie; et la faune mammalogique des couches à Pyrotherium. - Boletin del Instituto Geographico Argentino 15:501-660. - F. Ameghino - 1894.
- Mammiféres crétacés de l’Argentine (Deuxième contribution à la connaissance de la fauna mammalogique de couches à Pyrotherium) [Cretaceous mammals of Argentina (second contribution to the knowledge of the mammalian fauna of the Pyrotherium Beds)]. - Boletin Instituto Geografico Argentino 18(4–9):406-521. - F. Ameghino - 1897.
- Notices préliminaires sur des ongulés nouveaux des terrains crétacés de Patagonie [Preliminary notes on new ungulates from the Cretaceous terrains of Patagonia]. - Boletin de la Academia Nacional de Ciencias de Córdoba 16:349-429. - F. Ameghino - 1901.
- Pyrotherium macfaddeni, sp. nov. (late Oligocene, Bolivia) and the pedal morphology of pyrotheres. - Journal of Vertebrate Paleontology 24(2):481-488. - B. J. Shockey & F. Anaya-Daza - 2004.
- Enamel microstructure and mastication in Pyrotherium romeroi (Pyrotheria, Mammalia). Paläontologische Zeitschrift 89: 611–634. - W. Koenigswald, T. von Martin & G. Billet - 2015.



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