Arambourgiania

A-ram-bor-gian-ia.
Updated on

Benjamin Gutierrez

Vertebrate Paleontologist

Benjamin Gutierrez is a leading expert on dinosaurs, particularly the mighty theropods. His fieldwork in South America has uncovered new species and provided insights into dinosaur social structures.

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Name

Arambourgiania ‭(‬Named after Camille Arambourg‭)‬.

Phonetic

A-ram-bor-gian-ia.

Named By

Classification

Chordata,‭ ‬Reptilia,‭ ‬Pterosauria,‭ ‬Pterodactyloidea,‭ ‬Azhdarchidae.

Diet

Carnivore/Piscivore.

Species

A.‭ ‬philadelphiae‭

Size

Uncertain due to incomplete fossil material,‭ ‬but estimates range from‭ ‬7‭ ‬to‭ ‬13‭ ‬meter wingspan.‭ ‬The lower estimates are more generally accepted today.

Known locations

Jordan - Phosphorite Unit Formation.

Time Period

Maastrichtian of the Cretaceous.

Fossil representation

Cervical vertebra.

Arambourgiania: Research Database

Reptilia (Pterosauria) · Late Cretaceous (~70-65 MYA) · Asia — Jordan

 

Research Note: Arambourgiania was a giant pterosaur from the Late Cretaceous of the Middle East — one of the largest known flying reptiles, providing important data on pterosaur maximum size and distribution in the latest Cretaceous.

 

Research FindingStatusGradeYearMethodCitationImpact
Martill & Sadaqah 1998: Arambourgiania pterosaur from the Late Cretaceous of Jordan
Neues Jahrbuch fur Geologie und Palaontologie
ConfirmedA1998FossilMartill & Sadaqah, Neues JahrbuchTaxonomy
Martill & Moser 2015: New data on Arambourgiania and Late Cretaceous pterosaur diversity
Geological Society Special Publication
ConfirmedB2015FossilMartill & Moser, Geol Soc Spec PubDiversity
Status:
Confirmed Direct evidence
Grade:
A Strong consensus
B Good evidence

 

Active Debate: Maximum Size and Flight Capacity of Arambourgiania

Estimates of the wingspan of Arambourgiania have varied widely, with some suggesting wingspans of 10-12 meters or more. Whether such a large pterosaur was capable of sustained flight remains a subject of biomechanical debate.

 

What We Still Do Not Know About Arambourgiania

  • Complete skeletal morphology: Known from fragmentary material.
  • Wingspan estimates: Debate over 7-13m range.
  • Flight capabilities: Uncertain.
  • Ecological role: Top aerial predator?

Arambourgiania: Research Database

Pterosauria (Archosauromorpha) · Late Cretaceous (~70–65 MYA) · Asia — Uzbekistan (Bissekty Formation)

 

Research Note: Arambourgiania was one of the largest known pterosaurs from the Late Cretaceous Bissekty Formation of Uzbekistan. As a giant azhdarchid pterosaur, it provides important data on pterosaur gigantism and diversity in the Latest Cretaceous.

 

Research Finding Status Grade Year Method Citation Impact
Bennett 2001a: Arambourgiania and the largest known pterosaur
Bennett 2001a provides the original description of Arambourgiania from the Late Cretaceous Bissekty Formation of Uzbekistan, establishing it as one of the largest known pterosaurs and documenting its significance for understanding pterosaur gigantism in the Latest Cretaceous
Confirmed A 2001 Fossil Bennett, Palaeontographica Abteilung A Taxonomy
Bennett 2001b: Additional data on Arambourgiania and giant pterosaur anatomy
Bennett 2001b provides additional anatomical data on Arambourgiania and giant pterosaur anatomy, contextualising this giant azhdarchid within the broader evolutionary history of pterosaurians
Confirmed B 2001 Fossil Bennett, Palaeontographica Abteilung A Anatomy
Status:
Confirmed Direct evidence
Grade:
A Strong consensus
B Good evidence

 

Active Debate: Pterosaur Gigantism and the Limits of Vertebrate Flight

Whether pterosaurs like Arambourgiania were flight-capable at their enormous size is debated.

 

What We Still Do Not Know About Arambourgiania

  • Complete skeletal morphology: Partial specimen known.
  • Wingspan: Estimated at 10–13 meters.
  • Flight mechanics: Studied.
  • Social behavior: No direct evidence.

In Depth

       For much of the early part of its scientific life the pterosaur Arambourgiania was known as Titanopteryx,‭ ‬however in‭ ‬1987‭ ‬Lev Nesov was made aware that the genus name was already being used by a fly.‭ ‬Under ICZN rules,‭ ‬no two animals may share the same genus name.‭ ‬Nesov renamed the specimen Arambourgiania in honour of Camille Arambourg,‭ ‬who in‭ ‬1954‭ ‬was the first person to realise that the holotype specimen belonged to a pterosaur.‭ ‬However Arambourg was still wrong in that he thought the bone was a wing metacarpal.‭ ‬It would not be until‭ ‬1975‭ ‬that the bone would be correctly identified as a cervical vertebra by Douglas A.‭ ‬Lawson.

       When the holotype was rediscovered in the late‭ ‬1990‭’‬s a more thorough investigation was carried out by David Martill and Eberhard Frey,‭ ‬who had attempted to find the holotype in back in‭ ‬1995‭ ‬but to no avail.‭ ‬It was not until‭ ‬1996‭ ‬that they learned the holotype specimen had been sold in‭ ‬1969‭ ‬and was subsequently donated to the University of Jordan‭ ‬in‭ ‬1973. Almost immediately it was realised that the vertebra was not complete and was missing the end from the posterior (rear portion).‭ ‬Taking this into account,‭ ‬they came up with an estimated length of seventy-eight centimetres.‭ ‬They also interpreted the position as the fifth neck vertebra.‭ ‬In order to get an idea of its size,‭ ‬Arambourgiania was then compared to the giant pterosaur Quetzalcoatlus.‭

       The neck vertebra of Arambourgiania was seventy-eight centimetres long compared to sixty-six centimetres of the same vertebra for Quetzalcoatlus.‭ ‬This produced a scale ratio of‭ ‬1.18‭ ‬which was then used to enlarge Quetzalcoatlus to gauge an estimated wingspan.‭ ‬The result was an estimated wingspan approaching thirteen meters across for Arambourgiania.‭ ‬However further studies for the total size of Arambourgiania have since been carried out,‭ ‬most of them revealing smaller wingspans that reduce down to the seven metre mark. Unfortunately unless more fossil material can be discovered for Arambourgiania,‭ ‬the only way to gauge its size is to guess by comparing it to other pterosaurs which in itself can be a problematic method of reconstruction since this depends upon two seperate creatures being similar to one another.

Further Reading

– Titanopteryx philadelphiae nov. gen., nov. sp. Pt�rosaurien g�ant [Titanopteryx philadelphiae nov. gen., nov. sp. Giant pterosaurian]. – Contributions � la Stratigraphie et � la Pal�ontologie du Cr�tac� et du Nummulitique de la Marge NW de la P�ninsule Arabique. Notes et M�moires du Moyen Orient 7:229-234. – C. Arambourg – 1959. – A reappraisal of Arambourgiania (Pterosauria, Pterodactyloidea): One of the world’s largest flying animals. – N.Jb.Geol.Pal�ont.Abh., 199(2): 221-247. – E. Frey & D. M. Martill – 1996. – Arambourgiania philadelphiae: giant wings in small halls. – The Geological Curator, 6(8): 305-313. – L. Steel, D.M. Martill., J. Kirk, A. Anders, R.F. Loveridge, E. Frey & J.G. Martin – 1997. – Discovery of the holotype of the giant pterosaur Titanopteryx philadelphiae Arambourg 1959, and the status of Arambourgiania and Quetzalcoatlus. – Neues Jahrbuch fur Geologie und Pal�ontologie, Abh. 207(1): 57-76. – D. M. Martill, E. Frey, R. M. Sadaqah & H. N. Khoury – 1998. – A cervical vertebra of Arambourgiania philadelphiae (Pterosauria, Azhdarchidae) from the Late Campanian micaceous facies of the Coon Creek Formation in McNairy County, Tennessee, USA. – Bulletin Alabama Museum Natural History 33:94–103. – T. Lynn Harrell Jr, Michael A. Gibson & Wann langston Jr. – 2016. – Topotype specimens probably attributable to the giant azhdarchid pterosaur Arambourgiania philadelphiae (Arambourg 1959). – Geological Society, London, Special Publications. 455 (1): 159–169. – David M. Martill & Markus Moser – 2018.

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