Tropeognathus

Name: Tropeognathus ‭(‬Keel jaw‭)‬.
Phonetic: Trop-e-o-nay-fuss.
Named By: Peter Wellnhofer‭ ‬-‭ ‬1987.
Synonyms: Coloborhynchus mesembrinus,‭ ‬Criorhynchus mesembrinus,‭ ‬Ornithocheirus mesembrinus.
Classification: Chordata,‭ ‬Reptilia,‭ ‬Pterosauria,‭ ‬Ornithocheiridae.
Species: T.‭ ‬mesembrinus‭ (‬type‭)‬,‭ ‬T.‭ ‬robustus‭?
Diet: Piscivore.
Size: Largest known individual based upon specimen MN‭ ‬6594-1‭ ‬is estimated to have had a wingspan‭ ‬8.2‭ ‬meters across.
Known locations: Brazil‭ ‬-‭ ‬Santana Formation.
Time period: Aptian/Albian of the Cretaceous.
Fossil representation: Remains of at least three individuals,‭ ‬together revealing the skull and parts of the post cranial skeleton.

       Tropeognathus was first named in‭ ‬1987‭ ‬after the description of a pterosaur skull that had been purchased from a fossil dealer.‭ ‬In‭ ‬2002‭ ‬a set of lower jaws was further attributed to the genus,‭ ‬and most recently at the time of writing a third specimen was added in‭ ‬2013,‭ ‬and this was the first inclusion of post cranial remains.‭ ‬Today these specimens all represent members of the type species Tropeognathus mesembrinus which is held as valid,‭ ‬while a second species,‭ ‬Tropeognathus robustus which was also named in‭ ‬1987‭ ‬and based upon a more robust jaw,‭ ‬has a questionable validity.
       Only two years after Tropeognathus was established,‭ ‬there was speculation that it might have actually been a specimen of Anhanguera‭ (‬Kellner,‭ ‬1989‭)‬.‭ ‬Tropeognathus has also been considered a synonym to Coloborhynchus‭ (‬Veldmeijer,‭ ‬1998‭)‬,‭ ‬Criorhynchus‭ (‬Fastnacht,‭ ‬2001‭) ‬and Ornithocheirus‭ (‬Unwin,‭ ‬2001‭)‬.‭ ‬With the addition of new post cranial remains in‭ ‬2013‭ ‬however,‭ ‬Tropeognathus has now gained a wider acceptance to its validity,‭ ‬and a‭ ‬2013‭ ‬taxonomic review by Rodrigues and Kellner‭ ‬supported the view that Tropeognathus is a valid genus.
       Tropeognathus is easily amongst the largest known ornithocheirid pterosaurs,‭ ‬a kind that seem to have been best adapted for flying over bodies of water and snatching fish from the water.‭ ‬Also with the addition of new fossil material in‭ ‬2013,‭ ‬Tropeognathus was the largest pterosaur from Gondwana,‭ ‬the amalgamation of the southern continents that were largely separated from the‭ ‬northern continents that are known as Laurasia.‭ ‬Only the largest azhdarchid pteroasurs such as Quetzalcoatlus from North America are known to have comfortably exceeded Tropeognathus in size.‭ ‬Like relative genera,‭ ‬Tropeognathus‭ ‬had‭ ‬a toothed beak,‭ ‬and large rounded growths which have been dubbed‭ ‘‬keels‭’ ‬rising from the tips of both upper and lower jaws.‭
       The wings of Tropeognathus seem to be very long while at the same time thin,‭ ‬similar to some other types of pterosaurs such as pteranodonts.‭ ‬This may indicate that large ornithocheirds like Tropeognathus may have used a flight principal called dynamic soaring to conserve energy.‭ ‬This is where air current passing over ocean waves are exploited so that an individual can use them to pick up speed and lift without having to expend energy to continuously flap the‭ ‬wings.‭ ‬This idea of flight has also been proposed for the pterosaur Pteranodon,‭ ‬and can be seen in action by some modern sea birds such as the albatross‭ ‬which although has a different wing construction,‭ ‬has a roughly similar wing proportion.‭ ‬Glider pilots are also known to use this flight principal,‭ ‬though usually more often on a grander scale such as an air current rising over a mountain.

Further reading
-‭ ‬New crested pterosaurs from the Lower Cretaceous of Brazil.‭ ‬-‭ ‬Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historische Geologie‭ ‬27:‭ ‬175‭–‬186‭; ‬Muenchen‭ ‬-‭ ‬Peter Wellnhofer‭ ‬-‭ ‬1987.
-‭ ‬An overview of the pterosaur assemblage from the Cambridge Greensand‭ (‬Cretaceous‭) ‬of Eastern England.‭ ‬-‭ ‬Mitteilungen aus dem Museum für Naturkunde in Berlin,‭ ‬Geowissenschaftliche Reihe‭ ‬4:‭ ‬189‭–‬221.‭ ‬-‭ ‬D.‭ ‬M.‭ ‬Unwin‭ ‬-‭ ‬2001.
-‭ ‬First record of Coloborhynchus‭ (‬Pterosauria‭) ‬from the Santana Formation‭ (‬Lower Cretaceous‭) ‬of the Chapada do Araripe of Brazil.‭ ‬-‭ ‬Paläontologisches Zeitschrift,‭ ‬75:‭ ‬23‭–‬36.‭ ‬-‭ ‬M.‭ ‬Fastnacht‭ ‬-‭ ‬2001.
-‭ ‬Pterosaurs from the Lower Cretaceous of Brazil in the Stuttgart collection,‭ ‬-‭ ‬Stuttgarter Beiträge zur Naturkunde Serie B‭ (‬Geologie und Paläontologie‭) ‬327:‭ ‬1‭–‬27.‭ ‬-‭ ‬A.‭ ‬J.‭ ‬Veldmeijer‭ ‬-‭ ‬2002.
-‭ ‬The largest flying reptile from Gondwana:‭ ‬A new specimen of Tropeognathus cf.‭ ‬T.‭ mesembrinus Wellnhofer,‭ ‬1987‭ (‬Pterodactyloidea,‭ ‬Anhangueridae‭) ‬and other large pterosaurs from the Romualdo Formation,‭ ‬Lower Cretaceous,‭ ‬Brazil.‭ ‬Anais da Academia Brasileira de Ciências‭ ‬85:‭ ‬113.‭ ‬-‭ ‬A.‭ ‬W.‭ ‬A.‭ ‬Kellner,‭ ‬D.‭ ‬A.‭ ‬Campos,‭ ‬J.‭ ‬M.‭ ‬Sayão,‭ ‬A.‭ ‬N.‭ ‬A.‭ ‬F.‭ ‬Saraiva,‭ ‬T.‭ ‬Rodrigues,‭ ‬G.‭ ‬Oliveira,‭ ‬L.‭ ‬A.‭ ‬Cruz,‭ ‬F.‭ ‬R.‭ ‬Costa,‭ ‬H.‭ ‬P.‭ ‬Silva‭ & ‬J.‭ ‬S.‭ ‬Ferreira‭ ‬-‭ ‬2013.
-‭ ‬Taxonomic review of the Ornithocheirus complex‭ (‬Pterosauria‭) ‬from the Cretaceous of England.‭ ‬-‭ ‬ZooKeys‭ ‬308:‭ ‬1.‭ ‬-‭ ‬Taissa Rodrigues‭ & ‬Alexander Kellner‭ ‬-‭ ‬2013.



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