Name: Anhanguera ‭(‬Old devil‭).
Phonetic: Ahn-han-gair-ah.
Named By: Campos‭ & ‬Kellner‭ ‬-‭ ‬1985.
Synonyms: Araripesaurus santanae,‭ ‬Cearadactylus ligabuei,‭ ‬Ornithocheirus cuvieri,‭ ‬Pterodactylus fittoni,‭ ‬Santanadactylus araripensis,‭ ‬Tropeognathus robustus.
Classification: Chordata,‭ ‬Reptilia,‭ ‬Pterosauria,‭ ‬Pterodactyloidea,‭ ‬Ornithocheiridae,‭ ‬Anhanguerinae.
Species:‭ ‬‬A.‭ ‬blittersdorffi (type) A. piscator.
Diet: Piscivore.
Size:‭ U‬p to‭ ‬4.5‭ ‬meter wingspan.
Known locations: Australia.‭ ‬Brazil‭ ‬-‭ ‬Santana Formation.‭ ‬Russia.‭ ‬United Kingdom‭ ‬-‭ ‬Cambridge Greensand.
Time period: Aptian of the Cretaceous.
Fossil representation: Many specimens known.

       Like Ornithocheirus,‭ ‬Anhanguera had rounded crests on both the tip of its snout and lower jaw.‭ ‬Also the jaws broadened towards the tip with the sharp thin teeth pointing outwards in what is often referred to as a rosette.‭ ‬This arrangement greatly increased the‭ '‬catch area‭' ‬for Anhanguera as it flew over the water looking for fish. Anhanguera also had a smaller crest rising up from the back of its skull.‭ ‬This is quite a common feature amongst the advanced pterosaurs,‭ ‬though the crests of ornithocherid pterosaurs were not as dramatic as others such as Pteranodon and Nyctosaurus.
       With remains of Anhanguera being recovered in so many far reaching places,‭ ‬it seems that Anhanguera had a cosmopolitan distribution with the genus being represented‭ ‬by different species all over.‭ ‬However many of the species have since been found to represent other earlier named species.‭ ‬Anhanguera is also similar to the Chinese pterosaur Liaoningopterus.

Further reading
- Panorama of the flying reptiles study in Brazil and South America. - Anais da Academia Brasileira de Ciências 57(4):453-466. - D. de A. Campos & A. W. A. Kellner - 1985.
- Un novo exemplar de Anhanguera blittersdorffi (Reptilia, Pterosauria) da formaçao Santana, Cretaceo Inferior do Nordeste do Brasil [A new example of Anhanguera blittersdorffi (Reptilia, pterosaur) from the Santana Formation, Lower Cretaceous of Northeastern Brazil]. In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13. - - D. de A. Campos & A. W. A. Kellner - 1985.
- Description of a new species of Anhangueridae (Pterodactyloidea) with comments on the pterosaurfauna from the Santana Formation (Aptian–Albian), northeastern Brazil. - Tokyo, National Science Museum (National Science Museum Monographs, 17). - A.W.A. Kellner & Y. Tomida - 2000.
- Description of Coloborhynchus spielbergi sp. nov. (Pterodactyloidea) from the Albian (Lower Cretaceous) of Brazil. - Scripta Geologica, 125: 35-139. - A. J. Veldmeijer - 2003.
- Anhanguera taxonomy revisited: is our understanding of Santana Group pterosaur diversity biased by poor biological and stratigraphic control?. - PeerJ. 5: e3285. - F. L. Pinheiro & Taissa Rodrigues - 2017.


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